Asplenium trichomanes subsp. quadrivalens D.E.Mey., 1962

Common names

🇫🇷 Doradille fausse capillaire

Taxonomy

Kingdom: Plantae
Clade: Tracheophytes
Clade: Monilophytes
Class: Polypodiopsida
Subclass: Polypodiidae
Order: Polypodiales
Suborder: Aspleniineae
Family: Aspleniaceae
Genus: Asplenium
Species: Asplenium trichomanes subsp. quadrivalens

Presence

Present.

Conservation status

Red List (Metropolitan France): LC Least Concern

Notes

Subspecies with a brown-black rachis, whose fronds gradually become narrower, whose terminal segment is very reduced, and whose pinnae are elongated. The pinnae are inserted nearly perpendicularly on the rachis and are contiguous. Tetraploid (2n = 144). Sometimes elevated at species rank under the name A. quadrivalens (D.E.Mey.) Landolt, 2010.

Hybridizes with:

A. foreziense Le Grand ex Magnier, 1884 to produce A. × pagesii nothosubsp. pagesii Litard., 1910, P;
A. obovatum subsp. billotii (F.W.Schultz) O.Bolòs, Vigo, Masalles & Ninot, 1990 to produce A. × aran-tohanum Alejandre & M.J.Escal., 2005, P;
A. ruta-muraria L., 1753 to produce A. × clermontiae nothosubsp. clermontiae Syme, 1886, P;
A. septentrionale subsp. septentrionale (L.) Hoffm., 1796 to produce A. × alternifolium nothosubsp. heufleri (Reichardt) Aizpuru, Catalán & Salvo, 1985, P.
A. trichomanes subsp. coriaceifolium Rasbach, K.Rasbach, Reichst. & Bennert, 1990 to produce A. trichomanes nothosubsp. barreraense Rasbach, K.Rasbach, Reichst. & Bennert, 1990;
A. trichomanes subsp. hastatum (Christ) S.Jess., 1995 to produce A. trichomanes nothosubsp. lovisianum S.Jess., 1995, P;
A. trichomanes subsp. inexpectans Lovis, 1964 to produce A. trichomanes nothosubsp. lucanum Cubas, Rosselló & Pangua, 1989, P;
A. trichomanes subsp. pachyrachis (Christ) Lovis & Reichst., 1980 to produce A. trichomanes nothosubsp. staufferi Lovis & Reichst., 1985, P;
A. trichomanes subsp. trichomanes L., 1753 to produce A. trichomanes nothosubsp. lusaticum (D.E.Mey.) Lawalrée, 1978, P.

Photographs

img/asplenium-trichomanes-quadrivalens.webp — *Asplenium trichomanes* subsp. *quadrivalens*

img/asplenium-trichomanes-quadrivalens-2.webp — *Asplenium trichomanes* subsp. *quadrivalens*

img/asplenium-trichomanes-quadrivalens-3.webp — *Asplenium trichomanes* subsp. *quadrivalens*

img/asplenium-trichomanes-quadrivalens-4.webp — *Asplenium trichomanes* subsp. *quadrivalens*

img/asplenium-trichomanes-quadrivalens-5.webp — *Asplenium trichomanes* subsp. *quadrivalens* - Pinnae

img/asplenium-trichomanes-quadrivalens-6.webp — *Asplenium trichomanes* subsp. *quadrivalens* - Pinnae, abaxial side showing indusia

img/asplenium-trichomanes-quadrivalens-7.webp — *Asplenium trichomanes* subsp. *quadrivalens* - Pinnae, abaxial side showing indusia and sori

img/asplenium-trichomanes-quadrivalens-8.webp — *Asplenium trichomanes* subsp. *quadrivalens* - Pinnae, abaxial side showing open sporangia

img/asplenium-trichomanes-quadrivalens-9.webp — *Asplenium trichomanes* subsp. *quadrivalens* - Individual growing on the edge of a bridge

img/asplenium-trichomanes-quadrivalens-10.webp — *Asplenium trichomanes* subsp. *quadrivalens* - Individual growing on a wall

img/asplenium-trichomanes-quadrivalens-11.webp — *Asplenium trichomanes* subsp. *quadrivalens* - Individual growing on a wall

Sources

Badré, F., & Deschatres, R. (1979). Les Ptéridophytes de la France : liste commentée des espèces (taxinomie, cytologie, écologie et répartition générale). Candollea, 34(2), 379-457. https://doi.org/10.5169/seals-880127
Prelli, R., & Boudrie, M. (2024). Les fougères et plantes alliées d’Europe (2nd ed., p. 297). Biotope.
Tison, J.M., & de Foucault, B. (2014). Flora Gallica : Flore de France (p. 6). Biotope.