Polypodium × shivasiae Rothm., 1962

Common names

• 🇫🇷 Polypode de Shivas

Taxonomy

• Kingdom: Plantae
• Clade: Tracheophytes
• Clade: Monilophytes
• Class: Polypodiopsida
• Subclass: Polypodiidae
• Order: Polypodiales
• Suborder: Polypodiineae
• Family: Polypodiaceae
• Subfamily: Polypodioideae
• Genus: Polypodium
• Species: Polypodium × shivasiae

Presence

Present.

Notes

Hybrid of P. cambricum L., 1753 × P. interjectum Shivas, 1961.

The individual below was found in a mixed population of P. cambricum and P. interjectum. It is larger than the surrounding individuals of P. interjectum (hybrid vigor). It exhibits a morphology intermediate between its parent species: the blade shape is close to that of P. interjectum, the pinnae are pointed and the fronds bear a long terminal segment like P. cambricum. The sporangia are beige.

An examination under a microscope revealed a majority of aborted, grayish spores of variable size, and sporangia with an annulus made of 6-8 cells, with 2-3 cells between the pedicel and the base of the annulus.

Tetraploid (2n = 148).

Photographs

Sources

• Badré, F., & Deschatres, R. (1979). Les Ptéridophytes de la France : liste commentée des espèces (taxinomie, cytologie, écologie et répartition générale). Candollea, 34(2), 379-457. https://doi.org/10.5169/seals-880127

• Prelli, R., & Boudrie, M. (2024). Les fougères et plantes alliées d’Europe (2nd ed., p. 425). Biotope.

• Tison, J.M., & de Foucault, B. (2014). Flora Gallica : Flore de France (p. 23). Biotope.