Dryopteris affinis subsp. borreri (Newman) Fraser-Jenk., 1980
🇬🇧 Borrer's Male Fern
🇫🇷 Dryoptéride de Borrer, Dryoptéris de Borrer
Taxonomy
- Kingdom: Plantae
- Clade: Tracheophytes
- Clade: Monilophytes
- Class: Polypodiopsida
- Subclass: Polypodiidae
- Order: Polypodiales
- Suborder: Polypodiineae
- Family: Dryopteridaceae
- Subfamily: Dryopteridoideae
- Genus: Dryopteris
- Species: Dryopteris affinis subsp. borreri
Presence
Present.
Conservation status
- Red List (Metropolitan France): LC Least Concern
- Red List (Europe): LC Least Concern
Notes
Possible confusion with D. affinis subsp. affinis (Lowe) Fraser-Jenk., 1979, which has abundant reddish-brown scales on the petiole, and symmetrical basal pinnae with entire pinnules, while D. affinis subsp. borreri has few light brown scales on the petiole, more or less asymmetrical basal pinnae, and dentate or at least lobed basal pinnules.
This subspecies was previously treated at species level under the name D. borreri (Newman) Kinahan, 1856.
Apogamous triploid (n = 2n = 123).
The following infrasubspecific taxa are accepted:
- D. affinis subsp. borreri f. borreri (Newman) Fraser-Jenk., 1980, P;
- D. affinis subsp. borreri f. dilatata (Luerss.) Fraser-Jenk., 2024;
- D. affinis subsp. borreri f. robusta (Oberh. & Tavel ex Fraser-Jenk.) Fraser-Jenk., 2024.
Hybridizes with:
- D. filix-mas (L.) Schott, 1834 to produce D. × complexa nothosubsp. critica: pentaploid (2n = 205)
- nothof. critica Fraser-Jenk., 1987: f. borreri, P (hom. syn. D. × critica (Fraser-Jenk.) Fraser-Jenk., 2007);
- nothof. urbachensis (S.Jess. & Bujnoch) Fraser-Jenk., 2024: f. dilatata.
Named after William Borrer (1781-1862), an English botanist.
Photographs


































Sources
Badré, F., & Deschatres, R. (1979). Les Ptéridophytes de la France : liste commentée des espèces (taxinomie, cytologie, écologie et répartition générale). Candollea, 34(2), 379-457. https://doi.org/10.5169/seals-880127
Frasen-Jenkins, C.R. (2007). The species and subspecies in the Dryopteris affinis group. The Fern Gazette, 18(1), 1-26.
Prelli, R., & Boudrie, M. (2024). Les fougères et plantes alliées d’Europe (2nd ed., p. 365). Biotope.
Tison, J.M., & de Foucault, B. (2014). Flora Gallica : Flore de France (p. 12). Biotope.
Trewren, K. (2014). Some taxa within the Dryopteris affinis complex (p. 14). The British Pteridological Society.
Väre, H., Bento Elias, R., Christenhusz, M., Dyer, R., Ivanenko, Y., Rouhan, G., & Rumsey, F. (2017). Dryopteris borreri (Europe assessment). The IUCN Red List of Threatened Species, 2017. https://www.iucnredlist.org/species/83645436/84789468