Et ceterach…

Dryopteris affinis subsp. borreri (Newman) Fraser-Jenk., 1980

🇬🇧 Borrer's Male Fern

🇫🇷 Dryoptéride de Borrer, Dryoptéris de Borrer

Taxonomy

  • Kingdom: Plantae
  • Clade: Tracheophytes
  • Clade: Monilophytes
  • Class: Polypodiopsida
  • Subclass: Polypodiidae
  • Order: Polypodiales
  • Suborder: Polypodiineae
  • Family: Dryopteridaceae
  • Subfamily: Dryopteridoideae
  • Genus: Dryopteris
  • Species: Dryopteris affinis subsp. borreri

Presence

Present.

Conservation status

  • Red List (Metropolitan France): LC Least Concern
  • Red List (Europe): LC Least Concern

Notes

Possible confusion with D. affinis subsp. affinis (Lowe) Fraser-Jenk., 1979, which has abundant reddish-brown scales on the petiole, and symmetrical basal pinnae with entire pinnules, while D. affinis subsp. borreri has few light brown scales on the petiole, more or less asymmetrical basal pinnae, and dentate or at least lobed basal pinnules.

This subspecies was previously treated at species level under the name D. borreri (Newman) Kinahan, 1856.

Apogamous triploid (n = 2n = 123).

The following infrasubspecific taxa are accepted:

  • D. affinis subsp. borreri f. borreri (Newman) Fraser-Jenk., 1980, P;
  • D. affinis subsp. borreri f. dilatata (Luerss.) Fraser-Jenk., 2024;
  • D. affinis subsp. borreri f. robusta (Oberh. & Tavel ex Fraser-Jenk.) Fraser-Jenk., 2024.

Hybridizes with:

  • D. filix-mas (L.) Schott, 1834 to produce D. × complexa nothosubsp. critica: pentaploid (2n = 205)
    • nothof. critica Fraser-Jenk., 1987: f. borreri, P (hom. syn. D. × critica (Fraser-Jenk.) Fraser-Jenk., 2007);
    • nothof. urbachensis (S.Jess. & Bujnoch) Fraser-Jenk., 2024: f. dilatata.

Named after William Borrer (1781-1862), an English botanist.

Photographs

img/dryopteris-affinis-borreri.webp
Dryopteris affinis subsp. borreri
img/dryopteris-affinis-borreri-2.webp
Dryopteris affinis subsp. borreri
img/dryopteris-affinis-borreri-3.webp
Dryopteris affinis subsp. borreri - Pinnae
img/dryopteris-affinis-borreri-4.webp
Dryopteris affinis subsp. borreri - Pinnules
img/dryopteris-affinis-borreri-5.webp
Dryopteris affinis subsp. borreri - Pinnules, abaxial side showing indusia and sori
img/dryopteris-affinis-borreri-6.webp
Dryopteris affinis subsp. borreri - Basal pinnules of median pinnae
img/dryopteris-affinis-borreri-7.webp
Dryopteris affinis subsp. borreri - Basal pinnae
img/dryopteris-affinis-borreri-8.webp
Dryopteris affinis subsp. borreri - Basal pinnae
img/dryopteris-affinis-borreri-9.webp
Dryopteris affinis subsp. borreri - Black spots at the junction of the pinnae with the rachis
img/dryopteris-affinis-borreri-10.webp
Dryopteris affinis subsp. borreri - Petiole showing light brown scales
img/dryopteris-affinis-borreri-11.webp
Dryopteris affinis subsp. borreri - Croziers unfurling
img/dryopteris-affinis-borreri-12.webp
Dryopteris affinis subsp. borreri - Large root of an old individual, with a trunk-like appearance

img/dryopteris-affinis-borreri-13.webp
Dryopteris affinis subsp. borreri
img/dryopteris-affinis-borreri-14.webp
Dryopteris affinis subsp. borreri
img/dryopteris-affinis-borreri-15.webp
Dryopteris affinis subsp. borreri - Pinnae
img/dryopteris-affinis-borreri-16.webp
Dryopteris affinis subsp. borreri - Pinnules
img/dryopteris-affinis-borreri-17.webp
Dryopteris affinis subsp. borreri - Basal pinna
img/dryopteris-affinis-borreri-18.webp
Dryopteris affinis subsp. borreri - Basal pinnules of a basal pinna

img/dryopteris-affinis-borreri-19.webp
Dryopteris affinis subsp. borreri
img/dryopteris-affinis-borreri-20.webp
Dryopteris affinis subsp. borreri
img/dryopteris-affinis-borreri-21.webp
Dryopteris affinis subsp. borreri - Pinnae
img/dryopteris-affinis-borreri-22.webp
Dryopteris affinis subsp. borreri - Pinnules
img/dryopteris-affinis-borreri-23.webp
Dryopteris affinis subsp. borreri - Pinnules, abaxial side showing indusia
img/dryopteris-affinis-borreri-24.webp
Dryopteris affinis subsp. borreri - Basal pinnae
img/dryopteris-affinis-borreri-25.webp
Dryopteris affinis subsp. borreri - Black spots at the junction of the pinnae with the rachis
img/dryopteris-affinis-borreri-26.webp
Dryopteris affinis subsp. borreri - Petiole showing light brown scales

img/dryopteris-affinis-borreri-27.webp
Dryopteris affinis subsp. borreri
img/dryopteris-affinis-borreri-28.webp
Dryopteris affinis subsp. borreri
img/dryopteris-affinis-borreri-29.webp
Dryopteris affinis subsp. borreri - Pinnae
img/dryopteris-affinis-borreri-30.webp
Dryopteris affinis subsp. borreri - Pinnules
img/dryopteris-affinis-borreri-31.webp
Dryopteris affinis subsp. borreri - Basal pinnules of median pinnae
img/dryopteris-affinis-borreri-32.webp
Dryopteris affinis subsp. borreri - Basal pinnae
img/dryopteris-affinis-borreri-33.webp
Dryopteris affinis subsp. borreri - Black spots at the junction of the pinnae with the rachis
img/dryopteris-affinis-borreri-34.webp
Dryopteris affinis subsp. borreri - Petiole showing light brown scales

Sources

  • Badré, F., & Deschatres, R. (1979). Les Ptéridophytes de la France : liste commentée des espèces (taxinomie, cytologie, écologie et répartition générale). Candollea, 34(2), 379-457. https://doi.org/10.5169/seals-880127

  • Frasen-Jenkins, C.R. (2007). The species and subspecies in the Dryopteris affinis group. The Fern Gazette, 18(1), 1-26.

  • Prelli, R., & Boudrie, M. (2024). Les fougères et plantes alliées d’Europe (2nd ed., p. 365). Biotope.

  • Tison, J.M., & de Foucault, B. (2014). Flora Gallica : Flore de France (p. 12). Biotope.

  • Trewren, K. (2014). Some taxa within the Dryopteris affinis complex (p. 14). The British Pteridological Society.

  • Väre, H., Bento Elias, R., Christenhusz, M., Dyer, R., Ivanenko, Y., Rouhan, G., & Rumsey, F. (2017). Dryopteris borreri (Europe assessment). The IUCN Red List of Threatened Species, 2017. https://www.iucnredlist.org/species/83645436/84789468